Recent members

Dr Gustavo Martins (PhD student 2005-2009;  Postdoctoral collaborator 2009-2013 )


My interests are broad and span a wide range of topics but I am generally interested in describing and explaining the spatial (or temporal) distribution of marine organisms.

In 2009 I was awarded a post-doctoral grant (FCT, Portugal) where I examined how habitat isolation shapes community structure and affects ecosystem processes. This post-doctoral fellowship follows on from my PhD (2005-2009) which focused on the long-term community level impacts of exploitation of key intertidal herbivores (patellid limpets) on the Azorean archipelago.   

Theory predicts that fragmented and isolated habitats will on average support a less diverse biota. Patterns of species extinction in such habitats are, however, non-random with some species, usually generalists, being able to colonise and persist in isolated and small fragments, whilst larger and more specialised species are only able to persist in the larger and less isolated fragments. This suggests that functional traits are not evenly distributed in space but little is known as to how this will impact ecosystem processes. In my current research I use both observational and experimental work to seek a mechanistic link between isolation, community structure and ecosystem functioning.

Selected Publications:

Martins GM, Jenkins SR, Hawkins SJ, Neto AI, Thompson RC (in press) Illegal harvesting affects the success of fishing closure areas. Journal of the Marine Biological Association of the UK.

Jenkins SR, Martins GM (2010)Succession on Hard Substrata. In Dürr S, Thomason JC (eds) Biofouling, Wiley-Blackwell pp. 60-72

Martins GM, Thompson RC, Neto AI, Hawkins SJ, Jenkins SR (2010) Enhancing stocks of the exploited Patella candei via modifications in coastal engineering. Biological Conservation 143: 203-211.

Martins GM, Amaral AF, Wallenstein FM, Neto AI (2009) Influence of a breakwater on nearby rocky intertidal community structure. Marine Environmental Research 67:237-245.

Martins GM, Jenkins SR, Hawkins SJ, Neto AI, Thompson RC (2008) Exploitation of rocky intertidal grazers: population status and potential impacts on community structure and functioning. Aquatic Biology 3:1-10.

Martins GM, Thompson RC, Hawkins SJ, Neto AI, Jenkins SR (2008) Rocky intertidal community structure in oceanic islands: scales of spatial variation. Marine Ecology Progress Series 356: 15-24.

Martins GM, Hawkins SJ, Thompson RC, Jenkins SR (2007) Community structure and functioning in rock pools: effects of patch size and shore height at different successional stages. Marine Ecology Progress Series 329: 43-55.

Name: Dr Sergio Augusto Coelho de Souza (post-doc 2012-13)


I am interested in the ecology of ecosystems (functioning and biogeochemical cycles of carbon, nitrogen and mercury), in both marine and freshwater environments. I completed my masters in the Amazonian system focusing on mercury cycling and my PhD in a Brazilian upwelling system with pelagic microbial ecology as well as the interaction between bacteria and macroalgae in biofilms. My post-doc was in environmental education and scientific communication with traditional riverine communities in the Brazilian Amazon. In the School of Ocean Sciences (with a grant from Brazil, CNPq), I am studying the importance of biodiversity in both green and brown webs. 

Selected publications:

Coelho-Souza, SA; Miranda, MR; Salgado, LT; Coutinho, R; Guimaraes, JRD (2012) Adaptation of the [3]H-Leucine Incorporation Technique to Measure Heterotrophic Activity Associated with Biofilm on the Blades of the Seaweed Sargassum spp. Microbial Ecology. Accepted.

Coelho-Souza SA; López MS; Guimaraes JRD; Coutinho R; Candella RN (2012) Biophysical Interactions in the Cabo Frio Upwelling System, Southwest Brazil. Brazilian Journal of Oceanography. Accepted.

Curry JC; Araujo FV; Coelho-Souza SA; Peixoto RS; Olivera JAL; Santos HF; Davila AMR; Rosado AS (2011).Microbial Diversity of a Brazilian Coastal Region Influenced by an Upwelling System and Anthropogenic Activity. PLOS ONE 6 (1) e16553   DOI: 10.1371/journal.pone.0016553

Coelho-Souza, SA; Guimaraes, JRD; Miranda, MR; Poirier, H; Mauro, JBN; Lucotte, M; Mergler, D (2011).Mercury and flooding cycles in the Tapajos river basin, Brazilian Amazon: The role of periphyton of a floating macrophyte (Paspalum repens). Science of the total environment 409 (14): 2746- 2753   DOI:10.1016/j.scitotenv.2011.03.028

Coelho-Souza, SA; Guimaraes, JRD; Mauro, JBN; Miranda, MR; Azevedo, SMFO (2006). Mercury methylation and bacterial activity associated to tropical phytoplankton. Science of the total environment  364 (1-3):188 –199   DOI:10.1016/j.scitotenv.2005.07.010


Name: Dr María Soledad López (post-doc 2012-13)

I am broadly interested in ecological processes that shape rocky shore communities. During my PhD I investigated pre and post-settlement processes (larval supply, abiotic factors and prey-predator interactions) potentially responsible for the spatial and temporal distribution and abundance of a non-indigenous bivalve with a relatively recent history of introduction on the Brazilian coast. During my post-doc in Brazil I focused my work on prey phenotypic plasticity induced by the risk of predation and how some of these non-lethal predator effects can cascade to lower trophic levels in a simplified crab-gastropod-mussel trophic chain.

During my post-doctoral research in the UK (with a FAPESP grant, Brazil) I  extended my studies of non-lethal interactions by incorporating temporal variability and different levels of predation risk. For this purpose I explored the indirect non-lethal effect of Carcinus maenas on seaweed (Ascophyllum) traits mediated by changes in grazer (L. obtusata) activity.

 Selected Publications

Ignacio BL, López MS, Silva JSV (in press)  Colonization plasticity of the boring bivalve Lithophaga aristata (Dillwyn, 1817) on the Southeastern Brazilian coast: considerations on its invasiveness potential. Aquatic Invasions

López MS, Coutinho R, Ferreira CEL, Rilov G (2010) Predator- prey interactions in a bioinvasion scenario: differential predation by native predators on two exotic rocky intertidal bivalves. Marine Ecology Progress Series 403: 101-112.

López MS, Coutinho R (2010) Positive interaction between the native macroalgae Sargassum sp. and the exotic bivalve Isognomon bicolor. Brazilian Journal of Oceanography 58: 69-72.

Name Dr Gabriela Torres


I have general interests in biochemistry, ecophysiology and molecular biology, specially focussed in adaptations to stress. I was awarded my PhD by the University of Oldenburg, Germany. The research was conducted at the Helgoland Marine Station (AWI) Germany (DAAD scholarship) and at the University of Montpellier II in France (Socrates-Erasmus scholarship). My investigation was focussed on the ecophysiological adaptations to salinity variation in early life-history stages of decapod crustaceans, in particular growth, biochemical composition, enzymatic activity and mRNA expression of Na+-K+-ATPase.

During my postdoc at Bangor University, UK (Marie Curie Fellowship) I investigated the maternal and environmental effects on quality and performance of crustacean larvae. I used Carcinus maenas as model species and determined lipid profiles, elemental composition (CHN) and gene expression of transporter proteins: Na+-K-ATPase, Na+-K+-2Cl -contransporter and heat-shock proteins: HSP70, as well as survival and developmental rates as parameters.

From 2011 to 2014 I was involved in a project (NERC-Standard grant) to study the effects of larval quality on recruitment success using barnacles as model species (Austrominius modestus and Semibalanus balanoides). Since 2014  I have been working with German and Brazilian collaborators on a number of larval and zooplankton projects.

Selected publications:

Torres G, Giménez L, Anger K, 2011. Growth, tolerance to low salinity, and osmoregulation in decapod crustacean larvae. Aquatic Biology 12, 249-260.

Torres G, Giménez L, Anger K, 2008. Cumulative effects of low salinity on larval growth and biochemical composition in an estuarine crab, Neohelice granulata. Aquatic Biology 2, 37-45.

Anger K, Torres G, Nettelmann U, 2007. Adaptive traits in ecology, reproduction and early life history of Sesarma meridies, an endemic stream crab from Jamaica. Marine and Freshwater Research 58, 743-755.

Torres G, Charmantier-Daures M, Chifflet S, Anger K, 2007a. Effects of long-term exposure to different salinities on the location and activity of Na+-K+-ATPase in the gills of juvenile mitten crab, Eriocheir sinensis. Comparative Biochemistry and Physiology A 147, 460-465.

Torres G, Giménez L, Anger K, 2007b. Effects of osmotic stress on crustacean larval growth and protein and lipid levels are related to life-histories: the genus Armases as a model. Comparative Biochemistry and Physiology B, 148, 209- 224.

Anger K, Torres G, Giménez L, 2006. Metamorphosis of a sesarmid river crab, Armases roberti: stimulation by adult odours vs. inhibition by salinity stress. Marine Freshwater Behaviour and Physiology 39, 269-278.

Torres G, Anger K, Giménez L, 2006. Effects of reduced salinities on metamorphosis of a freshwater-tolerant sesarmid crab, Armases roberti: Is upstream migration in the megalopa stage constrained by increasing osmotic stress? Journal of Experimental Marine Biology and Ecology 338, 134-139.


Dr Kate Griffith

PhD student between 2009 and 2013 funded by a Bangor University 125 scholarship

Thesis title Climate driven changes in recruitment success of marine invertebrates: the role of food supply and temperature


In many marine invertebrates successful recruitment is dependent on synchronization of larval production with pulses of food (for example the spring phytoplankton bloom). A potentially important consequence of a warming climate is that shifts in larval phenology will cause larvae to be “mismatched” with their food sources. The extent to which marine invertebrates can respond to shifts in food availability is not clearly known but could lead to recruitment failure and population collapse of a species.  Kate's research investigated the role of food availability and temperature in the development and recruitment success of marine invertebrate larvae using  Semibalanus balanoides and Elminius modestus as model organisms.

Kate took up a post as a Lecturer in the School of Ocean Sciences in January 2014



Dr Timothy Whitton

PhD student between 2010 and 2014 funded by a University of Wales Studentship with sponsorship from the Environment Agency Wales

Thesis title: Processes controlling the spatial and temporal pattern in the cockle Cerastoderma edule


Tim's PhD focused on the common cockle Cerastoderma edule (L.), a commercially exploited bivalve throughout north western European shores. Work combined field surveys of settlement and post-settlement distribution with laboratory manipulations  of flow and predator interactions.  

Tim has taken up a position working within the project SEACAMS at the School of Ocean Sciences


Whitton T, Richardson CA, Jenkins SR, Hiddink J, 2012  The effect of prey aggregation on predation rates: juvenile shore crabs (Carcinus maenas) foraging on post-larval cockles (Cerastoderma edule).  Journal of Experimental Marine Biology and Ecology 432-433: 29-36 PDF

Dr Marija Sciberras

PhD student between 2008 and 2012 funded by the Maltese government, the IMAREST Stanley Gray Fellowship and the David Edward Roberts Memorial Award, Bangor University

Thesis title: Marine Protected Areas: Efficacy, Implementation and Management


Marija's PhD investigated the efficacy and ecological benefits generated by partially protected marine areas through two approaches:

1) a thorough examination of existing scientific and grey literature through a systematic review, to determine the relationship between the level of protection from anthropogenic extractive activities and the magnitude of the species’ response to protection

2)  collection of field data from a number of protected areas namely Port Erin dredging exclusion zone, Isle of Man; Skomer Marine Nature Reserve; ‘Modiolus box’, Llyn Peninsula; and Cardigan Bay Special Area of Conservation (SAC).

Following completion of her PhD Marija worked as a post-doc in Plymouth for one year before returning to Bangor to work with Jan Hiddink on an EU project examining the impacts of demersal fishing


Sciberras M, Hinz H, Bennell J, Jenkins SR, Hawkins SJ, Kaiser MJ 2013 Benthic community response to a scallop dredging closure within a dynamic seabed habitat.  Marine Ecology Progress Series 480:83-98 PDF

Sciberras M, Jenkins SR, Kaiser MJ, Hawkins SJ, Pullin AS 2013Evaluating the relative conservation value of partially and fully protected marine reserves.  Fish and Fisheries. DOI:  10.1111/faf.12044 PDF

Sciberras M, Jenkins SR, Kaiser MJ, Hawkins SJ and Pullin AS. 2013 Evaluating the biological effectiveness of fully and partially protected marine areas. Environmental Evidence 2:4 doi:10.1186/2047-2382-2-4


Dr Katrin Bohn 

PhD student between 2008 and 2012 funded by Countryside Council for Wales and Bangor Mussel Producers Organisation,

Thesis title: The distribution, spread, and impacts of the invasive marine gastropod Crepidula fornicata in Welsh waters 


Katrin's PhD focused on the distribution and spread of Crepidula fornicata in Wales, UK. This invasive marine gastropod was introduced to Europe from the North American Atlantic coast in the late 1870s and has since become a common component of the fauna along the east and south coast of England, where it may have major impacts on native species. On the west coast of the UK, however, it has not yet extended its range north of the Milford Haven Waterway in Wales, despite its rapid spread within the estuary and the high densities it reaches locally.   Through field and laboratory observation of larval supply/ settlement and  larval habitat selection combined with experimental work on limiting factors such as low temperature Katrin's PhD investigated the potential of this species to expand its range to mid and north Wales through natural larval dispersal. 

Following a post-doctoral position with Prof Steve Hawkins at Southampton working on (amongst other things) coastal protection through the THESEUS project, Katrin is now working on the Citizen Science project 'Capturing our Coast' at Portsmouth with Dr Gordon Watson.


Bohn K, Richardson CA, Jenkins SR 2013 Larval microhabitat associations of the non-native gastropod Crepidula fornicata and effects on recruitment success in the intertidal zone Journal of Experimental Marine Biology and Ecology 448: 289-297

Bohn K, Richardson CA, Jenkins SR 2013 The importance of larval supply, larval habitat selection and post-settlement mortality in determining intertidal adult abundance of the invasive gastropod Crepidula fornicata Journal of Experimental Marine Biology and Ecology 440: 132-140

Bohn K Richardson CA, Jenkins SR  2012The invasive gastropod Crepidula fornicata: reproduction and recruitment in the intertidal at its northern most range in Wales, U.K. and implications for its secondary spread.  Marine Biology 159: 2091-2103

Dr João Ferreira

PhD student between 2008 and 2012 funded by FCT (Fundação para a Ciência e a Tecnologia do Governo Português)

Thesis title: Latitudinal and climatic driven changes in local patterns of intertidal macroalgae: implications for biodiversity and ecosystem functioning 

Supervisors: Stuart Jenkins, Steve Hawkins (School of Ocean Sciences, Bangor University, Michael Burrows (Scottish Association for Marine Science, Oban, Scotland) 


From the Arctic to the Algarve there is a shift in the balance between macroalgae and grazers on intertidal rocky shores, with a change from macroalgal domination in the north to grazer domination in the south. Grazer control of macroalgae is thought to increase with declining latitude but the direct influence of climatic regime on early survival, growth and persistence of intertidal macroalgae is also important.   Patterns in intertidal community structure have been described qualitatively over these scales, but quantitative assessment of patterns of biomass and diversity and experiments to understand causal mechanisms are lacking.  In this context Joao's PhD  quantified rocky intertidal patterns over a European scale while exploring their mechanistic basis.

Joao took up a post-doc position in Stockholm, Sweden in 2014.


Ferreira JG, Arenas F, Martínez B, Hawkins SJ & Jenkins SR 2014 Physiological response of fucoid algae to environmental stress: comparing range centre and southern populations New Phytologist  In press

Ferreira JG, Hawkins SJ & Jenkins SR Patterns of reproductive output of fucoid species in core and marginal populations  Journal of Phycology In submission

Ferreira JG, Hawkins SJ & Jenkins SR  Physical and biological control of fucoid recruitment in range edge and range centre populations Marine Ecology Progress Series In submission

Dr Andrew Frederik Johnson 

PhD student between 2008 and 2012 funded by NERC with CASE partner Marine Biological Association of the UK

Thesis title: Determining the habitat requirements of demersal fish for the design of marine protected areas 

Supervisors: Stuart Jenkins, Hilmar Hinz, Jan Hiddink (School of Ocean Sciences, Bangor University); David Sims (Marine Biological Association of the UK, Plymouth)

Understanding the habitat requirements of key economic and ecologically important fish species is recognised as an integral part of  fishery management schemes.  Johnson's  PhD research (he prefers to be called Johnson for some strange reason) used descriptions of habitat (biotic and abiotic) and fish distribution data to elucidate factors determining the patterns found within fish communities at a range of spatial scales. He focused on the the strength of associations between demersal fishes and benthic habitat features such as sediment, prey abundance, biomass and size and in addition examined the effects of bottom trawling and how such anthropogenic impacts may alter such associations. Johnson's aim throughout was to increase understanding of benthos-fish linkages in order to help in the design and implementation of marine protected areas.

His work involved cruises in the Irish Sea (School of Ocean Sciences, Bangor), English Channel (Marine Biological Association, Plymouth) and Mediterranean (Instituto Espagnol de Oceanografia, Palma, Spain) using a range of techniques from GOC73 and beam bottom trawl surveys (sampling of fish and benthos) to grab sampling and underwater photography combined with multibeam sidescan bathymetry (descriptions of habitat). 

Since gaining his PhD in 2012 Johnson has travelled and worked in Chile (check out his blog ), Peru and India.

His work recently received attention in NERCs publication Planet Earth Online:

He now has a postdoc position at Scripps in San Diego, California.


Johnson AF, Gorrelli G, Jenkins SR, Hiddink JG, Hinz H, (in prep.) The impact of bottom fishing on the foraging of demersal fishes. Royal Society B: Biological Sciences.

Johnson AF, Jenkins SR, Hiddink JG, Hinz H, (2012) Linking temperate demersal fish species to habitat: scales, patterns and future directions. Fish and Fisheries. DOI: 10.1111/j.1467-2979.2012.00466.

Johnson AF, Valls Mir M, Moranta J, Jenkins S, Hiddink J, Hinz H, 2011. Effects of prey abundance and size on the distribution of demersal fishes.  Canadian Journal of Fisheries and Aquatic Sciences.

Hiddink JG, Johnson AF, Kingham R, Hinz H, Could our fisheries be more productive? Indirect negative effects of bottom trawl fisheries on fish condition. Journal of Applied Ecology. DOI: 10.1111/j.1365-2664.2011.02036.x

Griffiths AM, Sims DW, Johnson AF, Lynghammer A, Mchugh, M, Bakken T & Genner, MJ, 2011. Levels of connectivity between longnose skate (Dipturus oxyrinchus) in the Mediterranean Sea and the north-eastern Atlantic Ocean. Conservation Genetics. Doi:10.1007/s10592-010-0127-3

Griffin J, Mendez V, Johnson AF, Jenkins S, Foggo A, 2009. Functional Diversity Predicts Emergent Effects of Species Richness on Primary Productivity. Oikos doi: 10.1111/j.1600-0706.2008.16960.x

Johnson AF, Carew E, Sloman K, 2007. The Effects of Copper on the Morphological and Functional Development of Zebrafish Embryos. Aquatic Toxicology. doi:10.1016/j.aquatox.2007.07.003


Dr Ronaldo Adriano Christofoletti (post-doc 2008-2009)


Departamento de Ciências do Mar Universidade Federal de São Paulo Brazil

I am interested in the integration of ecological processes resulting in top-down or bottom-up control of community structure, depending on the relative influence of higher or lower trophic levels, and their spatial scale of influence. My work in the UK (Bangor University, May/08-Nov/09) focused on a simple predator-prey system (the predatory dogwhelk Nucella lapillus and filter feeders – barnacles and mussels) to evaluate the role of processes operating at a hierarchy of scales and particularly the importance of bottom-up control over large scales. This work was funded by a post-doctoral fellowship from CNPq – Conselho Nacional de Desenvolvimento Científico e Tecnológico, of the Ministry for Science and Technology of Brazil, under the supervision of Stuart Jenkins and collaboration of Michael Burrows.

At present, my principal interest is to understand the biotic control of ecosystem functioning and the negative impact of human activity on natural processes. Human activities such as harvesting, pollution, introduction of invasive species and, more recently, the impacts of climate change, can result in changes in the balance of natural processes (top-down and bottom-up). My work on rocky intertidal communities over the subtropical coast of Brazil addresses the relative importance of bottom-up (primary productivity, SST, organic matter, recruitment rates) and top-down control (diversity, abundance and size of consumers) in order to describe natural processes in this subtropical system and differences from temperate areas. Further workwill combine laboratory and field experimentationin an ecological model to determine how lateral modifiers (wave exposure, freshwater input, mangrove cover, coastline complexity), human impact (demography, contaminants in water, invasive species) as well as the impact of extreme environmental events (air temperature, humidity, rainfall) alter the balance of natural processes.

Selected Publications

Costa, T.M.; Christofoletti, R.A. & Pinheiro, M.A.A. 2010. Epibionts on Arenaeus cribrarius (Decapoda, Brachyura, Portunidae) from Brazil. Zoologia, 27 (3): 387-394.

Morishita, V.R.; Buchmann, F.S.C; Christofoletti, R.A.; Volpato, G. & Barreto, R.E. 2009. Prior residence and body size influence interactions between black sea urchins. Behavioral Processes, 80: 191-195.  

Sant’Anna, B.S; Christofoletti, R.A.; Zangrande, C.M. & Reigada, A.L.D. 2008. Growth of the hermit crab Clibanarius vittatus (Bosc, 1802) (Crustacea, Anomura, Diogenidae) at São Vicente, State of São Paulo, Brazil. Brazilian Archives of Biology and Technology, 51 (3): 547-550.

Madambashi, A.M.; Christofoletti, R.A. & Pinheiro, M.A.A. 2005. Natural diet of the crab Menippe nodifrons Stimpson, 1859 (Brachyura, Menippidae) in Parnapuã Beach, São Vicente (SP), Brazil. Nauplius 13 (1): 77-82.

Mantelatto, F.L.M.; Christofoletti, R.A. & Valenti, W.C. 2005. Population structure and growth of the hermit crab Pagurus brevidactylus (Stimpson, 1859) from Anchieta Island, São Paulo state, Brazil. Journal of the Marine Biological Association of United Kingdom, 85: 127-128.

Mantelatto, F.L.M.; R.A. Christofoletti & P.B. de Camargo. 2002. A food source analysis for the swimming crab Callinectes ornatus (Portunidae) in Ubatuba Bay (Brazil) using carbon isotopes. Nauplius, 10 (1): 61-65.

Mantelatto, F.L.M. & R.A. Christofoletti. 2001. Natural feeding activity of the crab Callinectes ornatus (Portunidae) in Ubatuba Bay (São Paulo, Brazil): influence of season, sex, size and moult stage. Marine Biology, 138 (3): 585-594.

Dr John Griffin

Contact: johngriffin@

John did his doctoral research under the supervision of Stuart Jenkins, Richard Thompson and Steve Hawkins at the Marine Biological Association of the UK and the University of Plymouth between 2005 and 2008. His work used rocky shores (and more specifically rock pools) as a model system to address links between aspects of biodiversity and ecosystem functioning. He focused on testing how environmental context can determine the impacts of species diversity (and conversely, extinction).

In 2009 John moved across the Atlantic to work as a postdoctoral associate at the University of Florida, with Brian Silliman to investigate the effects of predator diversity on the functioning of south eastern US salt marshes. Despite some Americanization, John stubbornly maintained his English accent, tried to drink at least one cup of tea a day, and returned to the UK in 2012 to take up a post as Lecturer at Swansea University.

Selected Publications

Griffin, J.N., Butler, J., Soomdat, N.N., Brun, K.E., Chejanovski, Z.A. & Silliman, B.R. (2011)    Top predators suppress rather than facilitate plants in a trait-mediated tri-trophic   cascade. Biology Letters, online early doi:10.1098/rsbl.2011.0166

Griffin, J.N., & Silliman, B.R. (2011) Predator diversity strengthens and stabilizes trophic  control of a keystone grazer. Biology Letters 7:79-82

Griffin, J.N., Noel, L., Hawkins, S.R., Thompson, R.C., & Jenkins S.J. (2010) Consumer effects on ecosystem functioning in rock pools: roles of species richness and composition.  Marine Ecology Progress Series 420: 45-56

Griffin, J.N., Jenkins, S.R., Hawkins, S.J., Gamfeldt, L. Thompson, R.C. (2009) Spatial heterogeneity increases the importance of species richness for an ecosystem process.  Oikos 118: 1335-1342

Griffin, J.N., Mendez, V., Johnson, A.F., Jenkins, S.R., Foggo, A. (2009) Functional diversity predicts overyielding effect of species combination on primary productivity.  Oikos: 118,  37-44

Griffin, J.N., de la Haye, K., Hawkins, S.J., Thompson, R.C., Jenkins S.R. (2008) Predator diversity and ecosystem functioning: density modifies the effect of resource partitioning. Ecology: 89(2), 298-305

Evelina Capasso

Evelina  spent five years working at the Marine Biological Association on long term change in benthic community structure, gaining her MPhil in 2011.  Her work was funded by Defra, Natural England, the MBA and for a period she was self funded.

Her work focused on decadal scale changes in the soft sediment benthic communities of the Western English Channel aiming  to contribute to the understanding of long-term changes occurring in marine ecosystems and specifically to separate natural and anthropogenic drivers. The approach chosen was not only at the community composition level, but through analysis of functional traits.  Evelina now works as an environmental consultant but continues to help with research at Bangor University on long term change.


Hinz H, Capasso E, Lilley M, Frost M, Jenkins SR  2011 Temporal differences across a bio-geographical boundary reveal slow response of sub-littoral benthos to climate change Marine Ecology Progress Series 423: 69-82

Capasso, E., Jenkins, SR., Frost, M., Hinz, H. (2010) Investigation of benthic community change over a century-wide scale in the Western English Channel. Journal of the Marine Biological Association of the UK 90(6):1161-1172

Dr Tom Potter

PhD sTom on the shore at Porth Cwyfantudent between 2011 and 2015 self-funded and by the Climate Change Consortium of Wales, IMAREST

Thesis title: The effects of warming on food web structure and function

Supervisors: Stuart Jenkins, Luis Gimenez, Andy Davies (School of Ocean Sciences, Bangor University)




Tom's research used a model consumer-producer system (the intertidal isopod Idotea granulosa and green alga Ulva lactuca) to understand how warming may modify the relationship between consumption and primary production.  He used both short and long term experiments in temperature controlled mesocosms to address these issues, as well as field obseravtions and manipulations. 

 Dr Kringpaka Wangkulangkul


PhD student between 2012 and 2016 funded by Prince of Songkla University, Thailand

Thesis title: Community level effects of variable recruitment of a key species in the rocky intertidal

Supervisors: Stuart Jenkins(School of Ocean Sciences, Bangor University) Steve Hawkins (University of Southampton)


The blue mussel Mytilus edulis is a common marine species found throughout Britain in a range of habitats, from subtidal sediments to intertidal rocky shores. Over recent decades, research has focused on the biology, physiology and fisheries aspect of M. edulis; however studies on its abundance and distribution on rocky shores and how these parameters influence the local community structure are scarce.  Kringpaka's research focused on


  • How the variability of mussel recruitment affects its abundance, distribution and the community structure on rocky shores over spatial and temporal scales
  • Mechanisms which potentially modify the relationship between recruitment and adult population structure
  • The mussel-fucoid interactions which potentially explain their distribution patterns on rocky shore